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The big thing with large-giant breed puppies is that they have to grow much faster than smaller puppies in a very similar amount of time to get to adult size. That fast growth can cause joint problems as the bones basically grow fast but not necessarily well. Limiting calories helps to limit the growth rate, and lower calorie (i.e., lower fat/protein) foods help to prevent over feeding.

They also need the proper amount of calcium: it should be between 0.8-1.1%. Too much calcium during puppyhood has been linked to skeletal disease later in life. (A lot of people talk about the P:Ca ratio but it's the total amount of Ca that appears to be the big problem, not necessarily it's ratio to P).
 
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