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Trey, my last sheltie, would heel very often, even up to old age. Honestly, the best thing for me was to stop it before it started. The behavior and desire to control motion was so strong in him that we never got it 100% fixed. A lot was observation and learning how to tell when he was over excited. He would bite when he was overstimulated and there would be signs leading up to the biting session.

A bite would mean all fun would stop. Turning away/ending things BEFORE the dog starts the biting is what I did/would do. Like I said, with observation there were signs leading up to it that would be a tell he was going to start trying to bite. Give him something productive to do then and to simultaneously get his head on a little clearer.

Honestly, though, your biting sounds a little different since he's gnawing. The herding dog type of biting I've experienced is usually a quick grip to the heel.
 
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